Table II

Mixed model analysis of several traits evaluated in two consecutive years (2006 and 2007): estimates (± SE) of phenotypic and genetic variances and family heritabilities. Traits evaluated were carbon isotope composition (δ¹³C), height increment (HI), diameter at breast height increment (DBHI), flushing date (Cf) , leaf fall date (LF) and growing season duration (GSD). Between brackets: probabilities associated with Wald Z-scores testing the null hypothesis of family variance component equal to zero. See text for explanation of variance component subscripts.

	Traits
Source	δ13C	HI	DBHI	Cf	LF	GSD
	2006
	0.23 ± 0.087 (0.003)	109.2 ± 56.66 (0.021)	1.40 ± 0.684 (0.021)	37.2 ± 11.94 (< 0.001)	29.7 ± 14.58 (0.021)	65.1 ± 26.54 (0.007)
	0.13 ± 0.029	0	0.22 ± 0.516	1.1 ± 0.89	24.4 ± 11.15	33.3 ± 13.51
	–	1214.6 ± 94.05	11.15 ± 0.918	14.9 ± 1.27	125.4 ± 10.77	122.8 ± 10.90
	0.84 ± 0.060	0.62 ± 0.126	0.67 ± 0.132	0.97 ± 0.012	0.66 ± 0.132	0.78 ± 0.086
	2007
	0.21 ± 0.084 (0.004)	34.9 ± 23.59 (0.068)	2.11 ± 1.192 (0.029)	32.6 ± 11.95 (0.003)	3.0 ± 2.78 (0.141)	46.0 ± 18.73 (0.009)
	0.18 ± 0.040	26.3 ± 26.18	1.88 ± 0.973	1.6 ± 2.23	7.8 ± 3.51	14.8 ± 6.11
	–	442.6 ± 37.94	12.73 ± 1.067	41.2 ± 3.55	42.3 ± 3.51	61.9 ± 5.35
	0.77 ± 0.086	0.51 ± 0.198	0.61 ± 0.159	0.92 ± 0.034	0.38 ± 0.246	0.85 ± 0.066