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Cited article:

Soluble nonprotein nitrogen compounds indicate changes in the nitrogen status of beech seedlings due to climate and thinning

Mariangela N. Fotelli, Michael Nahm, Anita Heidenfelder, Hans Papen, Heinz Rennenberg and Arthur Geßler
New Phytologist 154 (1) 85 (2002)
https://doi.org/10.1046/j.1469-8137.2002.00365.x